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|"We destroy. We rebuild. We destroy again. It is the constant of our kind. We are all a god's chosen, and we are all a god's cursed."|
|— Jon in Broken Throne|
Mare Barrow, Cal, Farley, and Shade first meet Jon when they are in the abandoned town of Rosen. He tells them that the newbloods of Pitarus, the location of their next recruitment mission, are dead and that Maven waits for them there.
Mare asks him to come with them, he refuses. However, he does help them before he leaves, telling them about Gareth and Nanny's imminent arrival, Julian and Sara's impending execution in Corros Prison, and Cameron's location. Mare trusted that he would be fine on his own and that he would be out of danger. Jon also tells them that their plan is going to work, but they have to leave instantly. When he's alone with Mare for a moment, he says to Mare that the answer to Farley's question is yes and that Mare will "rise alone." After this final warning, he leaves. It is later revealed that the question is whether or not Farley is pregnant with Shade's baby.
He later finds Maven and leads him on a path to intercept Mare's plane when she is flying to rescue the Dagger Legion, which leads to her capture.
Jon is serving as an advisor to King Maven. Mare sees him at Maven's side multiple times during her imprisonment at Whitefire Palace. At the banquet where the assassination attempt on Maven happens, Jon saves Mare's life by getting her attention. She turns to look at him, and the bullet that would of have killed her only grazes her. He then escapes from the Palace during the chaos.
In his first meeting with Mare, Jon reveals an outwardly calm personality, as he was able to brush off Farley pointing a gun at him with seemingly no fear due to the fact that he had looked into the future and knew he would not be shot. It is hinted that Jon holds a sadness within him due to his ability to "see everything," such as his own death. Despite his objections, it is shown that Jon likes using his ability to play games with people and enjoys the knowledge and power it grants him over other people. Jon never reveals what is in a person's future, but will sometimes hint at it. This makes him incredibly disliked by almost everyone. He is also very devious, considering he was able to betray both Mare, and later, Maven.
Jon has gray skin and long, gray hair. His most notable feature, however, are his blood red eyes. He wears gray clothes. Jon's nails are cracked and dirty. His skin is wrinkled despite his age. He appears bent with age, but has a feeling about him that causes others to fear him.
Jon has the mutation that grants him a Silver-like ability despite having Red blood. He has an ability similar to that of an eye, but is not limited to the immediate future. He has used this in conversations to answer people's questions before they can even ask them, as seen when he first met Mare, Cal, Farley, and Shade in Rosen. He also uses his ability to manipulate events and people, such as when he prevented Mare from walking into a trap Maven had set in Pitarus.
- Jon knew Shade was going to die at the raid on Corros Prison, but he did nothing to warn anyone because he also knew that Shade's death, and the raid, would save many people.